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Dr. William A. O’Donnell was born in Washington D.C. and grew up in Annandale, VA. He attended James Madison University, where he received a Bachelor of Science in Biology in 1977. He was on the faculty of Randolph Macon Academy, where he taught science, before attending the Medical College of Virginia School of Dentistry at Virginia Commonwealth University. He received his Doctorate of Dental Surgery in 1987.